TYPE: Costa Rica. San José: Pérez Zeledón, San Ramón Norte, trail to the summit of Cerro Pelón, 1420-1640 m, secondary mature and primary lower montane moist forest, 3 May 2000, F. Pupulin 2325, L. Spadari, J. Cambronero, V. Juárez-Pérez & K. Granado (holotype, JBL; illustration of type; photo of type).